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Puertasaurus reuili schematic. by randomdinos Puertasaurus reuili schematic. by randomdinos
Hold the door! Ever since its discovery in 2005, Puertasaurus has been claimed as one of the largest dinosaurs to ever exist, and, for once, although the downsize did happen, they weren't wrong. I was on a roll about the completeness of the animals while doing Ruyangosaurus and Patagotitan, which is... really not something that can be said for this genus, only having four vertebrae described and two measured and all that. Its inclusion within a clade with Patagotitan allowed me to mostly use the latter's proportions, although the position of Puerta as a lognkosaur is far from stable in their phylogeny, so its appearance may change eventually.

Puertasaurus is most remarkable for possessing the widest vertebrae of any known sauropod, and most likely of any animal ever discovered; scaling based on the ribcage of Patagotitan yields a torso around 3.6 meters wide, which may be less than the 5 meters originally estimated by Novas et al (2005), but would still be enough space to park a smart car sideways in - with room to spare. First thought to have been Maastrichtian in age, the Mata Amarilla (or Pari Aike) formation where Puertasaurus was found housed the megaraptoran Orkoraptor and the elasmarian Talenkauen.

PS: The mass discrepancies are of course uncertain - Puertasaurus has a shorter dorsal centrum and a far shorter overall vertebral height than Patagotitan, which indicates that its torso was not as deep, but the measurement across the diapophyses is 20% larger than in its relative, meaning it was, on the other hand, wider. The impact this has on the weight can vary from none to several tonnes depending on how much importance is given to diapophysis width; it's plausible that the mass difference between them could have been insignificant.

Previous version for comparison: sta.sh/01km6j7rrm22
First version: sta.sh/05w96zak9hx

References:
-
 Novas, Fernando E.; Salgado, Leonardo; Calvo, Jorge; Agnolin, Federico (2005). "Giant titanosaur (Dinosauria, Sauropoda)from the Late Cretaceous of Patagonia" (PDF)Revisto del Museo Argentino de Ciencias Naturales, n.s7 (1): 37–41. Archived from the original (PDF) on August 22, 2006. Retrieved 2007-03-04.
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 Gallina,P.A. and Apesteguía, S. 2011. Cranial anatomy and phylogenetic position of the titanosaurian sauropod Bonitasaura salgadoiActa Palaeontologica Polonica 56 (1): 45–60.
-Carballido, J.L.; Pol, D.; Otero, A.; Cerda, I.A.; Salgado, L.; Garrido, A.C.; Ramezani, J.; Cúneo, N.R.; Krause, J.M. (2017). "A new giant titanosaur sheds light on body mass evolution among sauropod dinosaurs". Proceedings of the Royal Society B: Biological Sciences284 (1860): 20171219. doi:10.1098/rspb.2017.1219.
-Gonzalez Riga, B.J., Mannion, P.D., Poropat, S.F., Ortiz David, L., Coria, J.P. 2018. Osteology of the Late Cretaceous Argentinean sauropod dinosaur Mendozasaurus neguyelap: implications for basal titanosaur relationships. Journal of the Linnean Society, zlx103, doi.org/10.1093/zoolinnean/zlx…


Human silhouette from www.onlygfx.com/20-woman-silho…
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:iconmark0731:
mark0731 Featured By Owner Edited Sep 4, 2018
Huh, it's shorter, but not/not really/barely lighter than I expected. I have no problem with it though, I still prefer it to the Scott Hartman version. So, it was likely more robust than Patagotitan and Argentinosaurus (absolutely compared to Patagotitan and proportionally compared to Argentinosaurus)?
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:iconrandomdinos:
randomdinos Featured By Owner Sep 4, 2018  Hobbyist Traditional Artist
I'm considering editing it to just be the same length as Patagotitan; the length of the centrum in Puertasaurus is estimated, and Patagotitan's C8 even more so.

Based on the only possible comparisons I'd say it had a proportionally wider but shallower back than both of them, even if the latter is only because of the neural arch (maybe the ribs were the same length). That would make the torso look more robust, but if it'd actually make it heavier per its total length isn't clear (of course, if it had a proportionally shorter neck, or shorter tail like with Hartman's, it would be heavier per TL)
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:iconmark0731:
mark0731 Featured By Owner Sep 4, 2018
"I'm considering editing it to just be the same length as Patagotitan"

Just because I wrote it's shorter than I expected, or you thought about it before that?
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:iconrandomdinos:
randomdinos Featured By Owner Edited Sep 4, 2018  Hobbyist Traditional Artist
Before. The D2 of Puertasaurus without the condyle is a couple centimeters longer than Patagotitan's in the same metric, but Patagotitan's is notably crushed, so the possibility of them being the same length was always a thing.
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:iconmark0731:
mark0731 Featured By Owner Edited Sep 4, 2018
Well, I like this version actually, but in the end, it's up to you. I guess the "Huh" at the beginning of my first sentence could've been misinterpreted as I was bummed because it came out to be shorter than Patagotitan, but I wasn't, I was just a bit surprised that it did.
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:iconmajestic-colossus:
Majestic-Colossus Featured By Owner Sep 3, 2018
Will Futa be updated too?
Reply
:iconrandomdinos:
randomdinos Featured By Owner Sep 3, 2018  Hobbyist Traditional Artist
It's definitely in need of one, I noticed that I didn't even upload a v2 of the schematic...
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:iconmajestic-colossus:
Majestic-Colossus Featured By Owner Sep 3, 2018
RIP, long, THICC tail. The rebirth of the 23m Futa is on the rise.
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:iconbac0nm0rph:
Bac0nm0rph Featured By Owner Sep 3, 2018  Hobbyist Artist
oh yes
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:iconjes86:
JES86 Featured By Owner Jun 6, 2018
These are all really well-done reconstructions.  I've only recently discovered your page - thanks to Franoys - but I'm starting to be a regular.
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:iconforbiddenparadise64:
ForbiddenParadise64 Featured By Owner Apr 20, 2018
Looks much better (not that the other looked bad or anything). I guess Puerta here is basically a somewhat chunkier version of Patagotitan (since its 4.6 tonnes heavier than Patagotitan would be scaled to the same length ) now? 
How does this affect the GDI statistics? I'm curious about them for...something I'm working on. 
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:iconrandomdinos:
randomdinos Featured By Owner Edited Apr 20, 2018  Hobbyist Traditional Artist
Even if the GDIs themselves changed (we're not going to re-GDI the lognkosaurs every time they're updated for... obvious reasons), the ratios should remain the same, since the discrepancies between them do. Although; the length estimate doesn't take into account that Patagotitan's d2 is more notably compressed than Puertasaurus's, yet both are the same length, so based on that comparison Puertasaurus should've had a shorter torso. This might've made them both the same size, which really wouldn't be unexpected.
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:iconforbiddenparadise64:
ForbiddenParadise64 Featured By Owner Apr 22, 2018
Puerta was indeed a thick one then...
Ah it's fine thanks. I was thinking of using a bit of your skeletals (with permission of course) for a fantasy project I have ideas for, as sauropod's are useful bases.
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:iconmark0731:
mark0731 Featured By Owner Oct 14, 2017
Damn! Titanosaurs, Y U NO reach 73 tonnes?
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:iconpaleosir:
paleosir Featured By Owner Oct 14, 2017  Hobbyist Traditional Artist
Considering the error bars still involved in reconstructing the giants, 73 tonnes is not out of the question.
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:iconmajestic-colossus:
Majestic-Colossus Featured By Owner Oct 14, 2017
Yep, ribcage width and shoulder height for example. The torso could still be taller (perhaps longer?) since we don't have any material from Puertasaurus itself to fill such gaps.
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:iconrandomdinos:
randomdinos Featured By Owner Edited Oct 14, 2017  Hobbyist Traditional Artist
(Or narrower and shorter, though I have yet to see anyone advocating for 37-tonne Puertasaurus ;) (Wink) ).
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:iconpaleosir:
paleosir Featured By Owner Oct 15, 2017  Hobbyist Traditional Artist
( I personally agree with your ~60 tonne estimate, I was just pointing out that there are still great error bars)
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:iconmajestic-colossus:
Majestic-Colossus Featured By Owner Oct 14, 2017
Yes, that's possible too!
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:iconpaleosir:
paleosir Featured By Owner Oct 14, 2017  Hobbyist Traditional Artist
Yes, and considering how different Puertasaurus dorsals look from Patagotitan´s , I doubt that using Patagotitan as a proxy for it´s size is terribly accurate (though this is not criticism for the skeletal: we simply don´t have much better).
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:iconfranoys:
Franoys Featured By Owner Edited Oct 14, 2017  Student Digital Artist
Puertasaurus D2 isn't all that much different from Patagotitan's in terms of proportions, both are of similar length once the condyle is excluded, in fact it is practically the same to the centimeter, the centrum width of Puertasaurus D2 is only 1 cm wider than that of Patagotitan's D2 (60 to 59 cm) and the centrum height vs centrum length discrepancy between Puertasaurus and Patagotitan (as I said the centrum length is roughly equivalent and the centrum height in Patagotitan is only a 7,5% bigger than that of Puertasaurus) is much smaller than between Patagotitan and  Futalognkosaurus (Puertasaurus D2 is less than a 5% longer than that of Futalognkosaurus but the centrum is 50% higher if we go by Hartman's skeletal of Futalognkosaurus to determine the centrum height).

Furthermore the D2 of Patagotitan is short compared to most of the dorsals in the dorsal series, so it is the animal that gets the longer torso length for Puertasaurus, in Futalognkosaurus D2-D10 are all about the same size. The only clear differences between Patagotitan and Puertasaurus are that in Puertasaurus each transverse process is about 14 cm longer and that Patagotitan has a bigger neural spine, but differences in this kind of prolongations are largely irrelevant when compared to the dimensions of the centra, which are the weight bearing portions.
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:iconmajestic-colossus:
Aside from the know materials of Patagotitan and Puertasaurus, we still don't really know that much about Puerta's shoulder height and limbs, right? Isn't that mostly speculation and use of other, better-known relatives?
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:iconfranoys:
Franoys Featured By Owner Oct 14, 2017  Student Digital Artist
Yes, the proportions of the limbs are entirely speculative, but it would be weird if they varied that much.
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:iconmajestic-colossus:
Majestic-Colossus Featured By Owner Oct 14, 2017
I agree, but who knows... xD
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:iconmajestic-colossus:
Majestic-Colossus Featured By Owner Oct 14, 2017
Yes. Probably not terribly accurate indeed, and I think nobody would claim that. Unfortunately, Puertasaurus is so incomplete that there's no other option left but using other species/genera to fill the gaps. Sadly that's the case with so many giant titanosaurs and sauropods in general. I hope more material gets unearthed.
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:iconpaleosir:
paleosir Featured By Owner Oct 14, 2017  Hobbyist Traditional Artist
We all hope, but considering the staggeringly low propabilities of even one Puertasaurus being preserved and found, I am not betting on it being found anytime soon (if at all).
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:iconmajestic-colossus:
Majestic-Colossus Featured By Owner Oct 14, 2017
It's already been more than a decade since Puertasaurus was discovered and so far nothing else was found (AFAIK). So yeah, that's sadly true. 
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:iconmark0731:
mark0731 Featured By Owner Oct 14, 2017
Well, yes, but with less probability :(
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:iconpaleosir:
paleosir Featured By Owner Oct 14, 2017  Hobbyist Traditional Artist
Maybe. There are quite a few differing opinions at the moment, though.
As Patagotitan showed, a random fairly complete new taxon can shake everything up.
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:iconevodolka:
Evodolka Featured By Owner Oct 14, 2017  Hobbyist Traditional Artist
i keep forgetting how little material we actually have on this guy :D
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:iconvasix:
vasix Featured By Owner Edited Oct 11, 2017  Hobbyist Digital Artist
With Bonitasaura being closer to lognkosaurs...might that make lognkosaurs, even ones of this size, giant, low-browsing animals? What are...your thoughts on that idea...? Especially considering Dreadnoughtus and Puertasaurus...coexisting? 
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:iconrandomdinos:
randomdinos Featured By Owner Oct 11, 2017  Hobbyist Traditional Artist
Heck... it might. Although I would find it hard to justify such an enormous neck and relatively tall shoulders if they were low browsers. Sounds like a waste of energy.

I'm guessing they could have been, at most, low-browsers relative to their size, but at a level that would still qualify as high-browsing by our standards; the shoulder of any of the three mega-lognkosaurs is already taller than the maximum head height of the largest giraffe ever recorded, after all.
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:iconlythronax-argestes:
lythronax-argestes Featured By Owner Oct 10, 2017  Hobbyist Digital Artist
HODOR
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:iconrandomdinos:
randomdinos Featured By Owner Edited Oct 11, 2017  Hobbyist Traditional Artist
It is known, that is how Puertasaurus vocalizations would have sounded
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:iconevodolka:
Evodolka Featured By Owner Oct 14, 2017  Hobbyist Traditional Artist
i can picture that
the young ones just shout "HOLD THE DOOR" and as the mature it gets less letters
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:icondontknowwhattodraw94:
Dontknowwhattodraw94 Featured By Owner Oct 10, 2017  Hobbyist Traditional Artist
That's big...
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:iconrandomdinos:
randomdinos Featured By Owner Edited Oct 10, 2017  Hobbyist Traditional Artist
Yes, but we can still go bigger. :D
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:icondontknowwhattodraw94:
Dontknowwhattodraw94 Featured By Owner Oct 11, 2017  Hobbyist Traditional Artist
Is that a hint for an upcoming Argentinosaurus? ;) (Wink) 
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:iconrandomdinos:
randomdinos Featured By Owner Oct 11, 2017  Hobbyist Traditional Artist
Soon enough. :P
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:iconpaleosir:
paleosir Featured By Owner Oct 10, 2017  Hobbyist Traditional Artist
Nice skeletal. 
But only 3m wide? Even Nima's Futalognkosaurus is wider than that.
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:iconmajestic-colossus:
Majestic-Colossus Featured By Owner Oct 10, 2017
He went more conservative about ribcage width, Nima apparently speculates more. But I was also expecting a pretty wide ribcage, possibly around 5m.
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:iconpaleosir:
paleosir Featured By Owner Oct 10, 2017  Hobbyist Traditional Artist
That is also what I expected.
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:iconrandomdinos:
randomdinos Featured By Owner Edited Oct 10, 2017  Hobbyist Traditional Artist
Thanks.

We have no indication of the width of Futalognkosaurus's ribcage more specific than "quite wide" and the few titanosaurs that do have measured ribcages aren't known for the uberwidth; Nima's is speculative, so the widths here and there can't be compared to one another. If you take a look at the ROM Futalognkosaurus, the width on that one is pretty similar to this proportionally.
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:iconpaleosir:
paleosir Featured By Owner Oct 10, 2017  Hobbyist Traditional Artist
I see. Thanks for the clarification.
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:iconrandomdinos:
randomdinos Featured By Owner Edited Oct 12, 2017  Hobbyist Traditional Artist
Actually, I was wrong - we do have an indication more specific than ''quite wide''. Futalognkosaurus's ilia are 255 wide across their widest spread point, which would make a width in the 3 m region more plausible for Patagotitan, and up to 3.6 m for Puertasaurus (3.35 m if we take the centrum width into account).
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:iconpaleosir:
paleosir Featured By Owner Oct 12, 2017  Hobbyist Traditional Artist
That makes more sense, because I already saw that the illia were very wide in Futalognkosaurus (But I hand't measured them or extrapolated them to Puertasaurus so I refrained from being specific)
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:iconkaprosuchusdragon:
KaprosuchusDragon Featured By Owner Oct 9, 2017  Hobbyist Digital Artist
:U
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:iconacepredator:
acepredator Featured By Owner Oct 9, 2017
Just do the damn French Monster
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